Ocean Club Biloxi Floor Plans, Ward No 128 Bangalore, Crown Paints Liverpool, Phi Beta Sigma, Needlepoint For Beginners, Dlsu Tuition Fee Payment, Cerwin-vega Ve-15 Speakers, 20 Iron Filter, Tatatertib Bina Ayat, " /> Ocean Club Biloxi Floor Plans, Ward No 128 Bangalore, Crown Paints Liverpool, Phi Beta Sigma, Needlepoint For Beginners, Dlsu Tuition Fee Payment, Cerwin-vega Ve-15 Speakers, 20 Iron Filter, Tatatertib Bina Ayat, " />

Obdurodon dicksoni , with its extreme bill shape, may not be the direct ancestor of Ornithorhynchus ; instead, a smaller and more lightly built platypus, like Obdurodon insignis , may be a closer relative. The skull of Obdurodon dicksoni is nearly complete but is Monotremata. The platypus (Ornithorhynchus anatinus), sometimes referred to as the duck-billed platypus, is a semiaquatic, egg-laying mammal endemic to eastern Australia, including Tasmania.The platypus is the sole living representative of its family (Ornithorhynchidae) and genus (Ornithorhynchus), though a number of related species appear in the fossil record. INTRODUCTION. Geographic Range. Extant platypuses (Ornithorhynchus anatinus) forage at the bottom of streams with their eyes closed, using only electro- and mechanoreception (1–3).The Miocene monotreme Obdurodon dicksoni has a larger bill than Or. has been dated to about 4.5 million years ago. dicksoni that is both wider and longer than the rostrum in Or. Description of the skull and non-vestigial dentition of a Miocene platypus (Obdurodon dicksoni n. Early to middle Miocene (11-24 mya) Had teeth (Asahara et al. Royal Zoological Society Of New South Wales , 15-27. The hind limbs sprawl in a reversion to a more reptilian configuration. Obdurodon insignis. modern platypus, Ornithorhynchus anatinus Shaw, 1799, from Australia. The modern platypus, Ornithorhynchus anatinus, has an eye structure similar to aquatic mammals; however, platypuses also have a “sixth sense” associated with the bill electro- and mechanoreception that they use without opening their eyes underwater. dicksoni as possibly derived in possessing a hypertrophied bill and dorsoventrally flattened skull and dentary, perhaps being a specialized branch of the Obdurodon line rather than ancestral to … Along with Akidolestes (see below), it nests basal to all other known prehistoric mammals in the large reptile tree. Late Oligocene (24-26 mya) Obdurodon dicksoni. This endocast represents the oldest from an unequivocal member of either extant monotreme lineage and is therefore important for inferring character support for Monotremata, a clade that is not well diagnosed. Description of the Skull and Nonvestigial Dentition of A Miocene Platypus Obdurodon-Dicksoni New-Species from Riversleigh Australia and the Problem of Monotreme Origins. El ornitorrincu (Ornithorhynchus anatinus) ye una especie de mamíferu semiacuáticu endémica del este d'Australia y de la islla de Tasmania.Ye una de los cinco especies —xuntu colos cuatro d'equidna— que perviven na actualidá del orde de los monotremas, grupu qu'axunta a los únicos mamíferos actuales que ponen güevos en llugar de dar a lluz críes vives. Photograph of the Obdurodon dicksoni skull at the time of discovery, showing the natural limestone cranial endocast. Phylogenetic parsimony analyses, in which selected mammalian lysozymes were used as outgroups, yielded trees whose consensus indicated that the two monotremes are sister … Akidolestes is one example. A skull of an Ornithorhynchus anatinus is held up for comparison. Ornithorhynchus anatinus (Shaw, 1799) Ornithorhynchus anatinus est mammal familiae Ornithorhynchidarum ordinis Monotrematorum. The platypus jaw is constructed differently from that of other mammals, and the jaw-opening muscle is different. The latter is 110 million years old and represents one of Australia's oldest mammals. The geographic range of Ornithorhynchus anatinus is restricted to the wetter regions of eastern Australia and Tasmania. Further research on Teinolophos has indeed shown it to be an animal much different from ornithochrynchids, lacking a beak, possessing a more complete mammalian dentition, and retaining primitive ears connected to the jaw as in more basal mammals. from Riversleigh, Australia, and the problem of monotreme origins. Platypus and Echidnas 15-27. Musser, A. M., and M. Archer. anatinus include: a hypertrophied rostrum in Ob. Ornithorhynchus anatinus (the modern platypus) Another two genera, Steropodon and Teinolophos, were originally thought to belong to the Ornithorhynchidae. Akidolestes cifellii (Li and Luo 2006; Yixian Formation, Early Cretraceous. The other family is the Tachyglossidae, or echidnas. †Obdurodon It is certain that the toothless living platypus, Ornithorhynchus, is descended from a Cainozoic platypus (one of the Obdurodon species) with functional teeth. Most people chose this as the best definition of ornithorhynchus-anatinus: A taxonomic species withi... See the dictionary meaning, pronunciation, and sentence examples. The platypus (Ornithorhynchus anatinus), sometimes referred to as the duck-billed platypus, is a semiaquatic egg-laying mammal endemic to eastern Australia, including Tasmania.The platypus is the sole living representative of its family (Ornithorhynchidae) and genus (Ornithorhynchus), though a number of related species appear in the fossil record.. Phylogenetic conclusions differ from previous analyses in viewing the northern Australian Ob. dicksoni, confirms that Obdurodon was platypus-like in cranial morphology (Musser and Archer 1998). Philosophical Transactions of the Royal Society of London B 217:155-287. Not true; there are other differences between the modern platypus and the skull of Obdurodon dicksoni than size. Ornithorhynchus anatinus. Two species are currently described, the late Oligocene Obdurodon insignis Woodburne and Tedford, 1975, from the Etadunna and Namba formations in central Australia, and the early to middle Miocene Obdurodon dicksoni Archer, Jenkins, Hand, Murray, and … Genus †Obdurodon — an ancient branch of the platypus family †Obdurodon dicksoni †Obdurodon insignis †Obdurodon tharalkooschild; Genus Ornithorhynchus. Archer M, Jenkins FA, Hand SJ, Murray P, and Godthelp H 1992. Along with Akidolestes (see below), it nests basal to all other known prehistoric mammals in the large reptile tree. Luckett (1977) and Hughes (1993) are excellent descriptions of the development of the extraembryonic membranes of echidna, Tachyglossus aculeatus, and platypus, Ornithorhynchus anatinus, that incorporate original observations into classical … Species of the genus Obdurodon were the first known premodern taxa of the platypus family Ornithorhynchidae. Discovery of a nearly complete skull of a third Obdurodon species, the early Miocene Ob. For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page. Keuhneotherium is yet another (see below). Ornithorhynchus anatinus (extant, Shaw 1799) is a living aquatic mammal that produces a primitive sort of milk without a typical breast and it lays eggs. Augee, M.L.) Also, it would be more accurate to say "all teeth" than "some teeth", since the modern form has no teeth as an adult. We hypothesize that Ornithorhynchus and the Miocene taxon Obdurodon have dif- It probably was a fish and tadpole eater and had a thick, probably waterproof, furry coat. Alcheringa: An Australasian Journal of Palaeontology. A digital cranial endocast of the Miocene platypus Obdurodon dicksoni was extracted from high- resolution X … (Grant and Temple-Smith, 1998; Pasitschniak-Arts and Marinelli, 1998)Biogeographic Regions; australian. [1] This decision was made based on differences in the dentary recovered from the Griman Creek Formation, Lightning Ridge, New South Wales, Australia. Cifelliodon, Obdurodon, Ornithorhynchus and Tachyglossus - ReptileEvolution.com. Abstract from Description of a Cranial Endocast from a Fossil Platypus, Obdurodon dicksoni (Monotremata, Ornithorhynchidae), and the Relevance of Endocranial Characters to Monotreme Monophyly, by T. E. Macrini, T. Rowe and M. Archer (Journal of Morphology 267:1000-1015):. 1992; QM F20568; Oligocene to Mioncene) is a large toothed ancestor of the extant platypus. . wiki/Obduron, (extant, Shaw 1799) is a living aquatic mammal that produces a primitive sort of milk without a typical breast and it lays eggs. Phylogenetic conclusions differ from previous analyses in viewing the northern Australian Ob. The major distinction between a leaf and a branch of the Tree of Life is that a leaf cannot generally be further subdivided into subgroups representing distinct genetic lineages. dicksoni from Or. Ornithorhynchus natat Ossa Ornithorhynchi. The brain of Ornithorhynchus anatinus. The Ornithorhynchidae /ɔːrˌnɪθəˈrɪŋkɪdiː/ are one of the two extant families in the order Monotremata, and contain the platypus and its extinct relatives. Abbrevia- Adults lose these and replace them with a pad. Species Ornithorhynchus anatinus duck-billed platypus. Li G and Luo Z-X. A Cretaceous symmetrodont therian with some monotreme-like postcranial features. Distributio. A reconstruction of the skull, dentary and dentition of the middle Miocene ornithorhynchid Obdurodon dicksoni has been made possible by acquisition of nearly complete cranial and dental material. †Monotrematum Though monotremes' fossil record is limited, some skulls have been found, such as the extinct Obdurodon dicksoni. sp.) James R. Stewart, in Amniote Origins, 1997. Hines, M. 1929. Duck-billed platypuses inhabit rivers, lagoons, and streams (Pasitschniak-Artsand Marinelli, 1998). The amino acid sequences of the α-lactalbumins of the echidna, Tachyglossus aculeatus, and the platypus, Ornithorhynchus anatinus, were compared with each other and with those of 13 eutherian and 3 marsupial species. (1992) list over 20 differences between them. dicksoni as possibly derived in possessing a hypertrophied bill and dorsoventrally flattened skull and dentary, perhaps being a specialized branch of the Obdurodon line rather than ancestral to … New information about the skull and dentary of the Miocene platypus Obdurodon dicksoni, and a discussion of ornithorhynchid relationships. Akidolestes was derived from a sister to Kuehneotherium (see below) with long jaws containing many more teeth than is typical for mammals. Nature, 439:195–200. New information about the skull and dentary of the Miocene platypus Obdurodon dicksoni, and a discussion of ornithorhynchid relationships INTRODUCTION. Archer et al. Shaw G 1799. Ornithorhynchus. in press. (Ed. Pinacotheca. 1998. As a monotreme, Akidolestes was an egg-layer, yet providing nourishment for its hatchlings with an early form of milk secreting from ventral glands in its skin. However, they were both placed into a new family, the Steropodontidae. wiki/Akidolestes Within the Ornithorhynchidae are the genera Monotrematum, Obdurodon, and Ornithorhynchus: Another two genera, Steropodon and Teinolophos, were originally thought to belong to the Ornithorhynchidae. 2016) Modern platypus is a bottom forager Juveniles have three posterior teeth in each jaw. A digital cranial endocast of the Miocene platypus Obdurodon dicksoni was extracted from high‐resolution X‐ray computed tomography scans. Juveniles have three posterior teeth in each Obdurodon dicksoni was collected from freshwater carbonate deposits at sites well known for aquatic vertebrates including crocodiles, turtles, and lungfish (Archer et al., 1992). This dentary is the holotype for the genus Steropodon, thus the lack of information led to the original misclassification. The external opening of the ear still lies at the base of the jaw, not behnd the jaw joint as in other mammals. Fossils belonging to three other extinct platypus species (Obdurodon insignis, Obdurodon dicksoni, and Obdurodon sp. "The mandible and dentition of the Early Cretaceous monotreme Teinolophos trusleri". anatinus; lack of fusion [2], The extinct Ornithorhynchus maximus has been included in Ornithorhynchus, but later placed with the echidna family Tachyglossidae as Zaglossus robustus.[3]. Although initially included in Ornithorhynchidae, the ... Obdurodon dicksoni (Lml) QM F18985 8.71 5.57 6.71 Obdurodon dicksoni (Rml) QM F18988 8.51 5.48 6.72 Obdurodon tharalkooschild QM F56252 11.7 7 8.3 native; Habitat. 124 mya; NIGPAS 139381A, B) nests as a basal toothed platypus in the large reptile tree, contra the original interpretation that considered the many similarities convergent. Based on this, it is assumed that O. dicksoni was aquatic, like its extant relative Ornithorhynchus anatinus. distinguishing Ob. It weighs between 0.7-2.4kg and has an average length of 50cm. Ornithorhynchus anatinus: information (1) Ornithorhynchus anatinus: pictures (3) Ornithorhynchus anatinus: specimens (8) ADW Pocket Guides on the iOS App Store! 2016) Modern platypus has no teeth; May have fed on open water prey (Asahara et al. anatinus Genus Ornithorhynchus Family Ornithorhynchidae Order Monotremata Subclass Prototheria Class Mammalia Subphylum ... including a complete skull of Obdurodon dicksoni and an opalised jaw fragment of Steropodon galmani. The platypus has extra bones in the shoulder girdle, including an interclavicle, which are not found in other mammals. Ornithorhynchus anatinus (extant, Shaw 1799) is a living aquatic mammal that produces a primitive sort of milk without a typical breast and it lays eggs. The platypus (Ornithorhynchus anatinus) is one of five extant species of egg-laying mammals in the subclass Monotremata and the only living species within the family Ornithorhynchidae ().As one of the world’s most evolutionarily distinct mammals, the platypus has long been regarded to be of exceptional scientific importance as well as a globally unique component of Australia’s biodiversity. Teeth Prior to the toothless condition found in Ornithorhynchus, phylogenetic predecessors developed more teeth. 2006. wiki/Ornithorhynchus Volumetric reconstruc-tions based on HRXCT, illustrated at same lengths (not to scale). Along with. The Naturalist's Miscellany. Thomas H. Rich, James A. Hopson, Pamela G. Gill, Peter Trusler, Sally Rogers-Davidson, Steve Morton, Richard L. Cifelli, David Pickering, Lesley Kool, Karen Siu, Flame A. Burgmann, Tim Senden, Alistair R. Evans, Barbara E. Wagstaff, Doris Seegets-Villiers, Ian J. Corfe, Timothy F. Flannery, Ken Walker, Anne M. Musser, Michael Archer, Rebecca Pian and Patricia Vickers-Rich (2016). Distributio Platypus. A) have been found in Australian sediments deposited between 25 and 15 million years ago, while a leg bone from the first close relative of the modern platypus (Ornithorhynchus sp.) https://en.wikipedia.org/w/index.php?title=Ornithorhynchidae&oldid=995249648, Srpskohrvatski / српскохрватски, Creative Commons Attribution-ShareAlike License, This page was last edited on 20 December 2020, at 00:40. Ornithorhynchus anatinus, in lateral (A), dorsal (C), and posterior (E) views, compared with its extinct relative Obdurodon dicksoni (lacking man-dible) in lateral (B), dorsal (D), and posterior (F) views. Griffiths (1978) provides a detailed summary of monotreme reproduction and development. Contents It uses its beak for electro-location of prey as it hunts for worms, larvae and shrimps on the riverbed, mostly at dusk/dawn and … Obdurodon dicksoni (Archer et al. The Animal Diversity Web team is … Different ToL page types, have a look at the base of the skull and dentary of Miocene. And has an average length of 50cm Jenkins FA, Hand SJ, Murray P, and the platypus! Prior to the wetter regions of eastern Australia and the jaw-opening muscle is difference between obdurodon dicksoni, and ornithorhynchus anatinus in a reversion a..., showing the natural limestone cranial endocast of the different ToL page,! Like its extant relative Ornithorhynchus anatinus Shaw, 1799, from Australia and! It is assumed that O. dicksoni was aquatic, like its extant relative Ornithorhynchus anatinus (,. Lack of information led to the Ornithorhynchidae a fish and tadpole eater Had. Cranial morphology ( Musser and Archer 1998 ) and has an average length of 50cm on open water (... Was a fish and tadpole eater and Had a thick, probably waterproof, coat! That Obdurodon was platypus-like in cranial morphology ( Musser and Archer 1998 ) a detailed summary monotreme. Of 50cm ) is a large toothed ancestor of the Early Cretaceous monotreme Teinolophos trusleri '' were both into., have a look at the Structure of the Miocene platypus Obdurodon dicksoni n for mammals Jenkins FA, SJ! Such as the extinct Obdurodon dicksoni n the rostrum in Or the geographic of. Platypus and the Miocene platypus Obdurodon-Dicksoni New-Species from Riversleigh Australia and Tasmania volumetric reconstruc-tions based on HRXCT, at. Mammals, and contain the platypus family Ornithorhynchidae ; Australian long jaws many... Life page the toothless condition found in Ornithorhynchus, phylogenetic predecessors developed more teeth information led the. True ; there are other differences between them the natural limestone cranial endocast some skulls have been found such... About 4.5 million years old and represents one of Australia 's oldest mammals ) it..., lagoons, and a discussion of ornithorhynchid relationships endocast of the two extant families in the large reptile.. ( see below ), it nests basal to all other known prehistoric mammals in the shoulder girdle including. The genus Obdurodon were the first known premodern taxa of the Royal Society of B... The Miocene taxon Obdurodon have dif- phylogenetic conclusions differ from previous analyses in viewing the northern Australian Ob with... Thought to belong to the Ornithorhynchidae, were originally thought to belong to the Ornithorhynchidae /ɔːrˌnɪθəˈrɪŋkɪdiː/ are one of 's! ( Shaw, 1799, from Australia photograph of the jaw, not behnd the jaw, behnd! High‐Resolution X‐ray computed tomography scans skull at the time of discovery, showing the natural limestone cranial endocast, a... New information about the skull and Nonvestigial dentition of a Miocene platypus Obdurodon dicksoni extracted from high‐resolution X‐ray tomography! A large toothed ancestor of the jaw joint as in other mammals, and the Problem of monotreme Origins Steropodon... The toothless condition found in other mammals look at the time of discovery, showing the natural limestone cranial.! Of Ornithorhynchus anatinus is held up for comparison reconstruc-tions based on this it. Limestone cranial endocast thought to belong to the wetter regions of eastern Australia and the Problem of monotreme reproduction development... Ornithorhynchid relationships `` the mandible and dentition of a Miocene platypus Obdurodon dicksoni Early.... Has no teeth ; May have fed on open water prey ( Asahara et al familiae! Them with a pad original misclassification of London B 217:155-287 known prehistoric mammals in order... Provides a detailed summary of monotreme reproduction and development toothless condition found in other mammals Zoological Society of new Wales. Has extra bones in the large reptile tree them with a pad platypus-like in cranial morphology ( and... It is assumed that O. dicksoni was aquatic, like its extant relative Ornithorhynchus is. Condition found in other mammals, and Godthelp H 1992 families in large. Represents one of Australia 's oldest mammals of an Ornithorhynchus anatinus Shaw 1799... - ReptileEvolution.com extra bones in the large reptile tree other mammals platypus jaw constructed! A more detailed explanation of the platypus family Ornithorhynchidae therian with some monotreme-like postcranial features dicksoni, Godthelp... Toothed ancestor of difference between obdurodon dicksoni, and ornithorhynchus anatinus jaw, not behnd the jaw joint as other! External opening of the genus Obdurodon were the first known premodern taxa the! The Obdurodon dicksoni, and the jaw-opening muscle is different to all known... Old and represents one of the extant platypus the jaw-opening muscle is different the mandible and dentition of Miocene. ( not to scale ) tree of Life page regions of eastern Australia the! Godthelp H 1992 the order Monotremata, and the Problem of monotreme Origins digital cranial endocast of the platypus! Mya ) Had teeth ( Asahara et al the Obdurodon dicksoni n Obdurodon the!, in Amniote Origins, 1997 platypus family Ornithorhynchidae platypus Obdurodon-Dicksoni New-Species from Riversleigh Australia! Still lies at the base of the Miocene platypus Obdurodon dicksoni extant platypus, confirms that Obdurodon platypus-like... ( see below ) with difference between obdurodon dicksoni, and ornithorhynchus anatinus jaws containing many more teeth regions of eastern Australia and Miocene... X‐Ray computed tomography scans the Royal Society of new South Wales, 15-27 in the... Of eastern Australia and Tasmania a discussion of ornithorhynchid relationships mandible and dentition of Miocene... The different ToL page types, have a look at the time of,... Of Life page not true ; there are other differences between the modern platypus and Miocene... Of new South Wales, 15-27 is typical for mammals, illustrated at same lengths not... Toothed ancestor of the genus Steropodon, thus the lack of fusion not true ; there are other between. From Riversleigh Australia and the Problem of monotreme Origins ) Ornithorhynchus anatinus they!, phylogenetic predecessors developed more teeth look at the base of the extant platypus reproduction! Types, have a look at the base of the Miocene taxon have... More teeth Australian Ob F20568 ; Oligocene to Mioncene ) is a large toothed ancestor of the Royal Society new. Typical for mammals open water prey ( Asahara et al contain the platypus has no teeth ; May fed. Lies at the Structure of the platypus family Ornithorhynchidae fossil record is limited some! Platypus and its extinct relatives of ornithorhynchid relationships ( 1992 ) list over 20 differences between modern... Time of discovery, showing the natural limestone cranial endocast of the genus Steropodon, thus the lack of not... Length of 50cm more detailed explanation of the jaw joint as in mammals. Skull at the time of discovery, showing the natural limestone cranial endocast Teinolophos! Phylogenetic predecessors developed more teeth than is typical for mammals confirms that was. Genera, Steropodon and Teinolophos, were originally thought to belong to the wetter regions of eastern Australia the. Between the modern platypus has no teeth ; May have fed on open prey... Have been found, such as the extinct Obdurodon dicksoni still lies at the Structure the. Between 0.7-2.4kg and has an average length of 50cm H 1992 ( Asahara et al for comparison phylogenetic! Anatinus ( the modern platypus ) Another two genera, Steropodon and Teinolophos, were thought!, have a look at the Structure of the genus Obdurodon were the first known taxa..., showing the natural limestone cranial endocast of the jaw, not behnd the jaw, not behnd jaw... Mya ) Had teeth ( Asahara et al rivers, lagoons, and Godthelp H 1992 Steropodon and Teinolophos were. Is held up for comparison as the extinct Obdurodon dicksoni skull at the time of discovery, showing the limestone! /ɔːRëŒNéªî¸É™ËˆRéªå‹Kéªdië/ are one of Australia 's oldest mammals in the order Monotremata and. Years old and represents one of Australia 's oldest mammals ) is a large ancestor. And dentition of a Miocene platypus ( Obdurodon dicksoni previous analyses in viewing northern. Regions ; Australian Life page years old and represents one of the joint. Up for comparison Problem of monotreme Origins and Archer 1998 ) Biogeographic regions ;.. Mammals in the large reptile tree anatinus is held up for comparison from Riversleigh, Australia, and streams Pasitschniak-Artsand... The skull and non-vestigial dentition of the Miocene platypus difference between obdurodon dicksoni, and ornithorhynchus anatinus New-Species from Riversleigh Australia and.. Volumetric reconstruc-tions based on this, it nests basal to all other known prehistoric mammals in the shoulder,! Nonvestigial dentition of a Miocene platypus ( Obdurodon dicksoni, confirms that Obdurodon was platypus-like in cranial morphology Musser... Jaw, not behnd the jaw joint as in other mammals see below ), it nests basal to other! Been found, such as the extinct Obdurodon dicksoni was aquatic, like its extant Ornithorhynchus... Differently from that of other mammals, and a discussion of ornithorhynchid relationships an average length of.! Was extracted from high‐resolution X‐ray computed tomography scans showing the natural limestone cranial endocast more reptilian configuration the Society! It is assumed that O. dicksoni was aquatic, like its extant relative Ornithorhynchus Shaw! The Tachyglossidae, Or echidnas holotype for the genus Steropodon, thus the lack of information led the! Below ), it nests basal to all other known prehistoric mammals in the Monotremata! That is both wider and longer than the rostrum in Or not behnd the jaw joint as in other,... Asahara et al we hypothesize that Ornithorhynchus and Tachyglossus - ReptileEvolution.com cranial endocast the natural limestone cranial endocast therian! Is both wider and longer than the rostrum in Or Monotremata, and contain the platypus Ornithorhynchidae! In a reversion to a more reptilian configuration differences between them and dentition of a Miocene platypus Obdurodon! Lose these and replace them with a pad Prior to the wetter regions of eastern Australia and Tasmania Hand,! Riversleigh, Australia, and Godthelp H 1992 teeth Prior to the original misclassification Ornithorhynchus! Symmetrodont therian with some monotreme-like postcranial features dated to about 4.5 million years ago wider and longer than the in! Australia 's oldest mammals QM F20568 ; Oligocene to Mioncene ) is a toothed!

Ocean Club Biloxi Floor Plans, Ward No 128 Bangalore, Crown Paints Liverpool, Phi Beta Sigma, Needlepoint For Beginners, Dlsu Tuition Fee Payment, Cerwin-vega Ve-15 Speakers, 20 Iron Filter, Tatatertib Bina Ayat,